Phymaturus

Taxonavigation

Taxonavigation: Pleurodonta
Superregnum: Eukaryota Cladus: Unikonta Cladus: Opisthokonta Cladus: Holozoa Regnum: Animalia Subregnum: Eumetazoa Cladus: ParaHoxozoa Cladus: Bilateria Cladus: Nephrozoa Superphylum: Deuterostomia Phylum: Chordata Cladus: Olfactores Subphylum: Vertebrata Infraphylum: Gnathostomata Megaclassis: Osteichthyes Cladus: Sarcopterygii Cladus: Rhipidistia Cladus: Tetrapodomorpha Cladus: Eotetrapodiformes Cladus: Elpistostegalia Superclassis: Tetrapoda Cladus: Reptiliomorpha Cladus: Amniota Cladus: Sauropsida Classis: Reptilia Cladus: Eureptilia Cladus: Romeriida Cladus: Diapsida Cladus: Neodiapsida Cladus: Sauria Cladus: Lepidosauromorpha Subclassis: Lepidosauria Ordo: Squamata Cladus: Unidentata, Episquamata Cladus: Toxicofera Subordo: Iguania Infraordo: Pleurodonta

Familia: Liolaemidae
Genus: Phymaturus
Species (52): P. aguanegra – P. aguedae – P. alicahuense – P. antofagastensis – P. bibronii – P. cacivioi – P. calcogaster – P. camilae – P. castillensis – P. ceii – P. chenqueniyen – P. curivilcun – P. darwini – P. delheyi – P. denotatus – P. desuetus – P. dorsimaculatus – P. etheridgei – P. extrilidus – P. felixi – P. fiambala – P. indistinctus – P. katenke – P. laurenti – P. loboi – P. mallimaccii – P. manuelae – P. maquinchao – P. maulense – P. nevadoi – P. niger – P. palluma – P. patagonicus – P. payuniae – P. punae – P. querque – P. rahuensis – P. robustus – P. roigorum – P. sinervoi – P. sitesi – P. somuncurensis – P. spurcus – P. tenebrosus – P. timi – P. tromen – P. verdugo – P. videlai – P. vociferator – P. williamsi – P. yachanana – P. zapalensis

Name

Phymaturus Gravenhorst, 1838: 749 [conserved name]

Type species: Lacerta palluma Molina, 1782, by monotypy.
Placed on the Official List of Generic Names in Zoology by Opinion 2118 (2005).

References

Primary references

Gravenhorst, J.L.C. 1838. Beiträge zur genauern Kenntniss einiger Eidechsengattungen. Nova Acta Physico-Medica Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum 18: 712–784. BHL
Etheridge, R. & Savage, J.M. 2003. Phymaturus Gravenhorst, 1837 and Lacerta palluma Molina, 1782 (currently Phymaturus palluma: Repilia, Sauria): proposed conservtion of usage of the names by designation of a neotype for Lacerta palluma Molina, 1782. Bulletin of Zoological Nomenclature 60(1): 38–41. BHL Reference page.
International Commission on Zoological Nomenclature 2005. Opinion 2118 (Case 3225): Phymaturus Gravenhorst, 1837 and Lacerta palluma Molina, 1782 (currently Phymaturus palluma; Reptilia, Sauria): usage of the names conserved by the designation of a neotype for Lacerta palluma Molina, 1782. Bulletin of Zoological Nomenclature 62(2): 116–117. Reference page.
Cei, J.M. & Scolaro, J.A. 2006. The neotype of the type species of the neotropical iguanian genus Phymaturus: a critical commentary on a recent opinion of the International Commission on Zoological Nomenclature. Zootaxa 1297(1): 17–22. DOI: 10.11646/zootaxa.1297.1.2 Reference page.
Etheridge, R. & Savage, J.M. 2006. Stability restored to the name Phymaturus palluma: a response to Cei and Scolaro (2006). Herpetological Review 37(4): 406–407. PDF (385-418). Reference page.

Additional references

Cei, J.M. & Castro, L.P. 1973. Taxonomic and serological researches on the Phymaturus patagonicus complex. Journal of Herpetology 7(3): 237–247. DOI: 10.2307/1563009 Reference page.
Cei, J.M. & Roig, V.G. 1975. A new lizard from the Sierra del Nevado Mountains, Central Argentina. Journal of Herpetology 9(2): 256. DOI: 10.2307/1563055 Reference page.
Etheridge, R. 1995. Redescription of Ctenoblepharys adspersa Tschudi, 1845, and the taxonomy of Liolaeminae (Reptilia: Squamata: Tropiduridae). American Museum Novitates 3142: 1–34. hdl: 2246/3658 Reference page.
Cei, J.M. & Videla, F. 2003. A new Phymaturus species from volcanic cordilleran mountains of the south-western Mendoza province, Argentina (Liolaemidae, Iguania, Lacertilia, Reptilia). Bollettino del Museo Regionale di Scienze Naturali di Torino 20(2): 291–314. hdl: 10915/85385 . Reference page.
Scolaro, J.A. & Cei, J.M. 2003. Una excepcional nueva especie de Phymaturus de la Precordillera de Chubut, Argentina (Liolaemidae, Iguania, Lacertilia, Reptilia) (In Spanish). Facena 19: 107–112. Full article (PDF). Reference page.
Pincheira-Donoso, D. 2004. Una nueva especie del género Phymaturus (Iguania: Tropiduridae: Liolaemini) del Centro-Sur de Chile (In Spanish). Multequina 13: 57–70. Full article (PDF). Reference page.
Lobo, F. & Quinteros, S. 2005. A morphology-based phylogeny of Phymaturus (Iguania: Liolaemidae) with the description of four new species from Argentina. Papéis Avulsos de Zoologia 45(13): 143–177. DOI: 10.1590/S0031-10492005001300001 Reference page.
Lobo, F. & Quinteros, S. 2005. Taxonomic studies of the genus Phymaturus (Iguania: Liolaemidae): Redescription of Phymaturus patagonicus Koslowsky 1898, and revalidation and redescription of Phymaturus spurcus Barbour 1921. Journal of Herpetology 39(4): 533–540. DOI: 10.1670/170-04A.1 Reference page.
Lobo, F. & Abdala, C.S. 2007. Descripción de una nueva especie de Phymaturus del grupo de P. palluma de la provincia de Mendoza, Argentina (In Spanish). Cuadernos de Herpetología 21(2): 103–113. Online Reference page.
Scolaro, J.A. & Ibargüengoytía, N.R. 2007. A new species of Phymaturus from rocky outcrops in the central steppe of Rio Negro province, Patagonia Argentina (Reptilia: Iguania: Liolaemidae). Zootaxa 1524(1): 47–55. DOI: 10.11646/zootaxa.1524.1.4 Reference page.
Scolaro, J.A. & Ibargüengoytía, N.R. 2008. A new fragment for the understanding of the puzzling evolutive process of the Phymaturus genus: a new species of the patagonicus group from Patagonia, Argentina (Reptilia: Iguania: Liolaemidae). Zootaxa 1939(1): 38–50. DOI: 10.11646/zootaxa.1939.1.4 Reference page.
Scolaro, J.A., Ibargüengoytía, N.R. & Pincheira-Donoso, D. 2008. When starvation challenges the tradition of niche conservatism: on a new species of the saxicolous genus Phymaturus from Patagonia Argentina with pseudoarboreal foraging behaviour (Iguania, Liolaemidae). Zootaxa 60(1): 48–60. DOI: 10.11646/zootaxa.1786.1.4 Reference page.
Scolaro, J.A. & Tappari, O.F. 2009. Una nueva especie del género Phymaturus del “grupo patagonicus” en los afloramientos rocosos del sudoeste de la provincia de Rio Negro, Patagonia Argentina (Reptilia: Iguania: Liolaemidae) (In Spanish). Naturalia Patagónica 4(2): 80–93. Full issue. Reference page.
Corbalán, V., Scolaro, J.A. & Debandi, G. 2009. A new species of the genus Phymaturus of the flagellifer group from central-western Mendoza, Argentina (Reptilia: Iguania: Liolaemidae). Zootaxa 2021(1): 42–56. DOI: 10.11646/zootaxa.2021.1.3 Reference page.
Scolaro, J.A. & Pincheira-Donoso, D. 2010. Lizards at the end of the world: two new species of Phymaturus of the patagonicus clade (Squamata, Liolaemidae) revealed in southern Patagonia of Argentina. Zootaxa 2393(1): 17–32. DOI: 10.11646/zootaxa.2393.1.2 Reference page.
Lobo, F., Abdala, C. & Valdecantos, S. 2010. Taxonomic Studies of the Genus Phymaturus (Iguania: Liolaemidae): Description of Four New Species. South American Journal of Herpetology 5(2): 102–127. DOI: 10.2994/057.005.0205 Reference page.
Núñez, H., Veloso, A., Espejo, P., Veloso, C., Cortés, A. & Araya, S. 2010. Nuevas especies de Phymaturus (grupo palluma) para la zona Cordillerana Central de Chile (Reptilia, Sauria, Liolaemidae) (In Spanish). Boletín del Museo Nacional de Historia Natural 59: 41–74. Full article (PDF). Reference page.
Ávila, L.J., Pérez, C.H.F., Pérez, D.R. & Morando, M. 2011. Two new mountain lizard species of the Phymaturus genus (Squamata: Iguania) from northwestern Patagonia, Argentina. Zootaxa 2924(1): 1–21. DOI: 10.11646/zootaxa.2924.1.1 Reference page.
Scolaro, J.A., Méndez-de la Cruz, F. & Ibargüengoytía, N.R. 2012. A new species of Phymaturus of the patagonicus clade (Squamata, Liolaemidae) from isolated plateau of southwestern Rio Negro Province, Argentina. Zootaxa 3451(1): 17–30. DOI: 10.11646/zootaxa.3451.1.2 Reference page.
Troncoso-Palacios, J. & Lobo, F. 2012. A new species of Phymaturus (Iguania: Liolaemidae) of the palluma group from Central Chile. Cuadernos de Herpetología 26(2): 69–78. Online. Reference page.
Lobo, F., Cruz, F.B., Abdala, C.S. 2012. Multiple lines of evidence show that Phymaturus agilis Scolaro, Ibargüengoytía & Pincheira-Donoso, 2008 is a junior synonym of Phymaturus spectabilis Lobo & Quinteros, 2005. Cuadernos de Herpetología 26(1): 21–27. Online. Reference page.
Lobo, F., Abdala, C. & Valdecantos, S. 2012. Morphological diversity and phylogenetic relationships within a South-American clade of iguanian lizards (Liolaemidae: Phymaturus). Zootaxa 3315(1): 1–41. DOI: 10.11646/zootaxa.3315.1.1 Reference page.
Lobo, F., nenda, S.J. & Slodki, D. 2012. A New Lizard of Phymaturus (Iguania: Liolaemidae) from Argentina. Herpetologica 68(1): 121–133. DOI: 10.1655/HERPETOLOGICA-D-11-00044.1 Reference page.
Lobo, F., Espinoza, R.E., Sanabria, E.A. & Quiroga, L.B. 2012. A New Phymaturus (Iguania: Liolaemidae) from the Southern Extreme of the Argentine Puna. Copeia 2012(1): 12–22. DOI: 10.1643/CH-11-086 Reference page.
Lobo, F., Laspiur, A. & Acosta, J.C. 2013. Description of new andean species of the genus Phymaturus (Iguania: Liolaemidae) from Northwestern Argentina. Zootaxa 3683(2): 117–132. DOI: 10.11646/zootaxa.3683.2.2 Reference page.
Scolaro, J.A., Jara, M. & Pincheira-Donoso, D. 2013. The sexual signals of speciation? A new sexually dimorphic Phymaturus species of the patagonicus clade from Patagonia Argentina. Zootaxa 3722(3): 317–332. DOI: 10.11646/zootaxa.3722.3.2 Reference page.
Morando, M., Ávila, L.J., Pérez, C.H.F., Hawkins, M.A. & Sites Jr., J.W. 2013. A molecular phylogeny of the lizard genus Phymaturus (Squamata, Liolaemini): implications for species diversity and historical biogeography of southern South America. Molecular Phylogenetics and Evolution 66(3): 694–714. DOI: 10.1016/j.ympev.2012.10.019 . ResearchGate . Reference page.
Troncoso-Palacios, J., Lobo, F., Etheridge, R., Acosta, J.C. & Laspiur, A. 2013. The rediscovery of Oplurus bibronii Guichenot, 1848 a valid species of the liolaemid genus Phymaturus (Iguania: Liolaemidae). Zootaxa 3652(5): 547–561. DOI: 10.11646/zootaxa.3652.5.4 Reference page.
Ávila, L.J., Pérez, C.H.F., Minoli, I. & Morando, M. 2014. A new lizard of the Phymaturus genus (Squamata: Liolaemidae) from Sierra Grande, northeastern Patagonia, Argentina. Zootaxa 3793(1): 99–118. DOI: 10.11646/zootaxa.3793.1.4 Reference page.
Troncoso-Palacios, J. & Esquerré, D. 2014. A new species of Phymaturus of the P. mallimaccii Group from the Andes of central Chile (Iguania: Liolaemidae). Phyllomedusa 13(1): 3–15. DOI: 10.11606/issn.2316-9079.v13i1p3-15 Reference page.
Lobo, F. & Nenda, S.J. 2015. Discovery of two new species of Phymaturus (Iguania: Liolaemidae) from Patagonia, Argentina, and occurrence of melanism in the patagonicus group. Cuadernos de herpetología 29(1): 5–25. Abstract and full article (PDF). Reference page.
Corbalán, V., Debandi, G., Scolaro, J.A. & Ojeda, A.A. 2016. DNA Barcoding of Phymaturus Lizards Reveals Conflicts in Species Delimitation within the patagonicus Clade. Journal of Herpetology 50(4): 654–666. DOI: 10.1670/15-104 Reference page.
Lobo, F., Barrasso, D.A., Hibbard, T. & Basso, N.G. 2016. On the evolution and diversification of an andean clade of reptiles: combining morphology and DNA sequences of the palluma group (Liolaemidae: Phymaturus). Zoological Journal of the Linnean Society 176(3): 648–673. DOI: 10.1111/zoj.12335 Reference page.
Marín, A.G., Pérez, C.H.F., Minoli, I., Morando, M. & Ávila, L.J. 2016. A new lizard species of the Phymaturus patagonicus group (Squamata: Liolaemini) from northern Patagonia, Neuquén, Argentina. Zootaxa 4121(4): 412–430. DOI: 10.11646/zootaxa.4121.4.3. Reference page.
Scolaro, J.A., Corbalán, V., Tappari, O.F. & Streitenberger, L.O. 2016. Lizards at the end of the world: a new melanic species of Phymaturus of the patagonicus clade from rocky outcrops in the northwestern steppe of Chubut province, Patagonia Argentina (Reptilia: Iguania: Liolaemidae). Boletín del Museo Nacional de Historia Natural 65: 137–152. Full article (PDF). Reference page.
Lobo, F., Barrasso, D.A., Paz, M. & Basso, N.G. 2018. Phylogenetic relationships within a Patagonian clade of reptiles (Liolaemidae: Phymaturus) based on DNA sequences and morphology. Journal of Zoological Systematics and Evolutionary Research 56(4): 549–569. DOI: 10.1111/jzs.12221 Reference page.
Troncoso-Palacios, J., Ferri-Yáñez, F., Laspiur, A. & Aguilar, C. 2018. An updated phylogeny and morphological study of the Phymaturus vociferator clade (Iguania: Liolaemidae). Zootaxa 4441(3): 447–466. DOI: 10.11646/zootaxa.4441.3.2 Reference page.
Hibbard, T.N., Nenda, S.J. & Lobo, F. 2019. A New Species of Phymaturus (Squamata: Liolaemidae) from the Auca Mahuida Natural Protected Area, Neuquén, Argentina, Based on Morphological and DNA Evidence. South American Journal of Herpetology 14(2): 123–135. DOI: 10.2994/SAJH-D-17-00067.1 Reference page.
Becker, L.A., Boretto, J.M., Cabezas-Cartes, F., Márquez, S., Kubisch, E., Scolaro, J.A., Sinervo, B. & Ibargüengoytía, N.R. 2019. An integrative approach to elucidate the taxonomic status of five species of Phymaturus Gravenhorst, 1837 (Squamata: Liolaemidae) from Northwestern Patagonia, Argentina. Zoological Journal of the Linnean Society 185(1): 268–282. DOI: 10.1093/zoolinnean/zly043 Reference page.
Lobo, F., Hibbard, T., Quipildor, M. & Valdecantos, S. 2019. A New Species of Lizard Endemic to Sierra de Fiambalá, Northwestern Argentina (Iguania: Liolaemidae: Phymaturus). Integrated Taxonomy Using Morphology and DNA Sequences: Reporting Variation Within the antofagastensis Lineage. Zoological Studies 58: 20. DOI: 10.6620/ZS.2019.58-20 Reference page.
Lobo, F., Barrasso, D.A., Hibbard, T., Quipildor, M., Slodki, D., Valdecantos, S. & Basso, N.G. 2021. Morphological and genetic divergence within the Phymaturus payuniae clade (Iguania: Liolaemidae), with the description of two new species. South American Journal of Herpetology 20(1): 42–66. DOI: 10.2994/SAJH-D-18-00082.1 Reference page.
Scolaro, J.A., Corbalán, V., Obregón Streitenberger, L. & Tappari, O.F. 2021. Description of Phymaturus katenke, a new species of lizard (Iguania: Liolaemidae) discovered through DNA barcoding. North-Western Journal of Zoology 17(1): 57–64. PDF Reference page.
Lobo, F., Barrasso, D.A., Valdecantos, S., Giraudo, A.R., Di Pietro, D.O. & Basso, N.G. 2022. Exploring the morphological diversity of Patagonian clades of Phymaturus (Iguania: Liolaemidae). Integrative study and the description of two new species. Cuadernos de Herpetología 36(2): 197–231. DOI: 10.31017/CdH.2022.(2022-015) Reference page.

Links

Uetz, P., Freed, P., Aguilar, P., Reyes, R., Kudera, F. & Hošek, J. (eds.) 2025. Phymaturus . The Reptile Database. Accessed on 26 May 2025.

Vernacular names

English: Rocky Lizards
español: Matuastos