When you're playing birds you can swap two food tokens for any other food token but does that also apply to activated abilities? eg.
Could I play two worm tokens in lieu of a fish token?
From the rulebook, page 6:
When playing birds, you may spend any 2 food tokens as if they are any 1 food token. This exchange cannot be used during other parts of the game.
So the answer is no - you can only substitute food tokens when playing birds, not when using other abilities.