Keith Richard Willmott

Keith Richard Willmott, English entomologist. Author abbreviation: Willmott

McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida, USA

Taxon names authored

(List may be incomplete)

129 taxon names authored by Keith Richard Willmott

Eponyms

(List may be incomplete)

1 eponym of Keith Richard Willmott

Publications

(List may be incomplete)

1994

Willmott, K.R. & Hall, J.P.W. 1994. Four new species of Riodinids from western Ecuador (Lepidoptera: Riodinidae). Tropical lepidoptera 5(2): 87-91. Full PDF. Reference page.

1995

Hall, J.P.W. & Willmott, K.R., 1995. Two new species of Adelpha from eastern Ecuador (Lepidoptera: Nymphalidae). Tropical Lepidoptera 6 (2): 106–109. Full article: . Reference page.
Hall, J.P.W. & Willmott, K.R. 1995a: Notes on the genus Argyrogrammana, with descriptions of five new species (Lepidoptera: Riodinidae). Tropical lepidoptera 6(2): 136–143. Full article (pdf). Reference page.
Hall, J.P.W. & K.R. Willmott, 1995b. Five new species and a new genus of riodinid from the cloud forests of eastern Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera Research 6(2): 131–135. Full article: PDF. Reference page.
Hall, J.P.W. & Willmott, K.R. 1995c. Two new species of Mesene from western Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera 6(2): 110-112. PDF. Reference page. .
Hall, J.P.W. & Willmott, K.R. 1995d. Two new species of satyrines from Ecuador (Nymphalidae: Satyrinae). Tropical Lepidoptera 6(2): 103-105. PDF. Reference page.

1996

Hall, J.P.W. & Willmott, K.R. 1996. Systematics of the riodinid tribe Symmachiini, with the description of a new genus and five new species from Ecuador, Venezuela and Brazil (Lepidoptera: Riodinidae). Lambillionea 96(4): 637–660. PDFa PDFb Reference page.
Hall, J.P.W. & Willmott, K.R. 1996. The genus Theope: four new species and a new subspecies (Lepidoptera: Riodinidae). Tropical lepidoptera 7(1): 63–67. Abstract (pdf). Reference page.
Hall, J.P.W. & Willmott, K.R., 1996. Notes on the genus Argyrogrammana. Part 2, with one new species (Lepidoptera: Riodinidae). Tropical Lepidoptera 7 (1): 71–80. Abstract: . Reference page.
Torres-Núñez, R., Hall, J.P.W., Willmott, K.R. & Johnson, K. 1996. A new genus of Elfin butterflies from the northern high Andes (Lepidoptera: Lycaenidae). Tropical lepidoptera 7(1):81-86.Reference page.

1998

Hall, J.P.W.; Willmott, K.R. 1998: Four new Riodinid species from eastern Ecuador (Lepidoptera: Riodinidae). Lambillionea 98(3): 325–334. Full article Reference page.
Hall, J.P.W. & Willmott, K.R. 1998. Nine new species and one new subspecies of Euselasia from Ecuador (Lepidoptera: Riodinidae). Tropical lepidoptera 9 supplement 1: 27–35. full article (pdf). Reference page.
Hall, J.P.W. & Willmott, K.R. 1998b. Three new species of Riodinini from the cloud forests of Ecuador. (Lepidoptera: Riodinidae). Tropical lepidoptera 9 supplement 1: 22–26. Full article (pdf). Reference page.

1999

Pyrcz, T.W., Willmott, K.R. & Hall, J.P.W., 1999: Contribution to the knowledge of Ecuadorian Pronophilini, Part III. Three new species and five new subspecies of Lymanopoda (Lepidoptera: Nymphalidae: Satyrinae). Genus 10(3): 497-522. full article (pdf)Reference page.

2001

Willmott, K.R., Hall, J.P.W. & Lamas, G. 2001. Systematics of Hypanartia (Lepidoptera: Nymphalidae: Nymphalinae), with a test for geographical speciation mechanisms in the Andes. Systematic Entomology 26: 369–399. PDF Reference page.

2004

Willmott, K.R. & Mallet, J. 2004. Correlations between adult mimicry and larval hostplants in ithomiine butterflies. Proceedings of the Royal Society of London B (Biology Letters Suppl.) 271: S266-S269. Reference page.

2005

Hall, J.P.W.; Willmott, K.R., Busby, R.C. 2005: Five new Penaincisalia species (Lepidoptera: Lycaenidae: Eumaeini) from the Andes of southern Ecuador and northern Peru. Zootaxa 797: 1–20. Abstract & excerpt Reference page.
Hall, J. & Willmott, 2005. A New Species of Paiwarria (Lepidoptera: Lycaenidae: Eumaeini) from Western Ecuador. Proceedings of the Entomological Society of Washington 107 (4): 960–967. BHL. Reference page.

2006

Willmott, K.R. & Freitas, A.V.L. 2006. Higher-level phylogeny of the Ithomiinae (Lepidoptera: Nymphalidae): classification, patterns of larval hostplant colonisation and diversification. Cladistics 22: 297-368. Reference page.
Willmott, K.R. & Lamas, G. 2006. A phylogenetic reassessment of Hyalenna Forbes and Dircenna Doubleday, with a revision of Hyalenna (Lepidoptera: Nymphalidae: Ithomiinae). Systematic Entomology 31(3): 419-468. Reference page.

2007

Hall & Willmott, [2007]. Four new species of Symmachiini from Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera 16 (1-2): 1–5. PDF. Reference page.
Willmott, K.R. & Lamas, G. 2007. A revision of Pachacutia, a new genus of rare Andean ithomiine butterflies (Nymphalidae: Ithomiinae), with the description of two new species. Annals of the Entomological Society of America 100: 449-469. DOI: 10.1603/0013-8746(2007)100[449:AROPAN]2.0.CO;2. . Reference page.

2008

Willmott, K.R. & Vitale, F. 2008. Taxonomic notes on Napeogenes (Lepidoptera: Ithomiinae) from Ecuador and Colombia, with the description of ten new subspecies. Genus 19(3): 585-609. Reference page.
Willmott, K.R. & Lamas, G. 2008. A revision of the genus Megoleria (Lepidoptera: Nymphalidae: Ithomiini, (Ithomiina), Tropical Lepidoptera Research 18(1): 36-59 (PDF). Reference page.

2011

Willmott, K.R., Elias, M. & Sourakov, A. 2011. Two Possible Caterpillar Mimicry Complexes in Neotropical Danaine Butterflies (Lepidoptera: Nymphalidae). Annals of the Entomological Society of America 104(6): 1108-1118. DOI: 10.1603/AN10086. . Reference page. .

2015

Nakahara, S., Hall, J.P.W., Lamas, G. & Willmott, K.R. 2015. Seven new species and one new subspecies of Euptychia Hübner, 1818 (Lepidoptera: Nymphalidae: Satyrinae) from the tropical Andes. Tropical Lepidoptera Research 25(2): 63–79. Reference page.
Willmott, K.R. 2015. A new subspecies of Cithaerias pyropina (Salvin & Godman, 1868) from the Cordillera de Cutucú in southeastern Ecuador (Nymphalidae: Satyrinae). Tropical lepidoptera Research 25(1): 1-4. full article (pdf). Reference page.

2016

Freitas, A.V.L., Barbosa, E.P., Willmott, K.R., Wahlberg, N. & Lamas, G. 2016. ‘Species’ from two different butterfly genera combined into one: description of a new genus of Euptychiina (Nymphalidae: Satyrinae) with unusually variable wing pattern. Revista Brasileire de Entomologia 60(3): 157–165. full article (PDF). Reference page.
Freitas, A.V.L., Magaldi, L.M. & Willmott, K.R. 2016. A new subspecies of Oleria gunilla (Nymphalidae: Danainae) from North Mato Grosso, Brazil. Journal of the Lepidopterists’ Society 70(4): 295–301 DOI: 10.18473/lepi.70i4.a8. Reference page.
Nakahara, S., Tan, D., Lamas, G., Parus, A. & Willmott, K.R. 2016. A distinctive new species of Hermeuptychia Forster, 1964 from the eastern tropical Andes (Lepidoptera: Nymphalidae: Satyrinae) Tropical Lepidoptera Research 26(1): 6-12. full article (PDF). Reference page.
Nakahara, S., Barbosa, E.P., Marin, M.A., Freitas, A.V.L., Pomerantz, T. & Willmott, K.R. 2016a. Graphita gen. nov., a New Genus for Neonympha griphe C. Felder & R. Felder, 1867. (Lepidoptera, Nymphalidae, Satyrinae). Neotropical Entomology 45: 675–691. DOI: 10.1007/s13744-016-0414-x. . Reference page.
Nakahara, S., Vega, G. & Willmott, K.R. 2016. Description of a new species of Euptychia Hübner, 1818 (Lepidoptera: Nymphalidae: Satyrinae) from the western Andes. Zootaxa 4184(2): 358–366. DOI: 10.11646/zootaxa.4184.2.7. Reference page.
Willmott, K.R., Mielke, O.H.H. & Freitas, A.V.L. 2016. A new subspecies and validation of the enigmatic species Adelpha herbita Weymer endemic to the Brazilian Atlantic forest (Nymphalidae: Limenitidinae). Tropical Lepidoptera Research 26(1): 32-37. full article (PDF). Reference page.

2017

Willmott, K.R., Lamas, G. & Hall, J.P.W. 2017. Notes on the taxonomy of Actinote intensa Jordan (Lepidoptera: Nymphalidae: Heliconiinae) and the description of a new sibling species from eastern Ecuador. Tropical Lepidoptera Research 27(1): 6-15. full article (PDF). Reference page.
Willmott, K.R., Robinson Willmott, J.C., Elias, M. & Jiggins, C.D. 2017a. Maintaining mimicry diversity: optimal warning color patterns differ among microhabitats in Amazonian clearwing butterflies. Proceedings of the Royal Society B 284(1855)(20170744): 1-9. DOI: 10.1098/rspb.2017.0744. . Reference page.
Willmott, K.R., Simon, M., Ortiz-Acevedo, E. & Hall, J.P.W. 2017. First record of the enigmatic tribe Anaeomorphini (Lepidoptera, Nymphalidae, Charaxinae) outside of the Amazon basin: a new species of Anaeomorpha Rothschild, 1894, from the Chocó region of western Ecuador. Insecta Mundi (0548): 1–10. Full article (PDF). Reference page.

2018

Barbosa, E.P., Siewert, R.R., Mielke, O.H.H., Lamas, G., Willmott, K.R. & Freitas, A.V.L.. 2018. Redescription of Yphthimoides patricia (Hayward, 1957), with taxonomic notes on the names Euptychia saltuensis Hayward, 1962 and Yphthimoides manasses (C. Felder & R. Felder, 1867) (Nymphalidae: Satyrinae). Zootaxa 4422(4): 537–557. DOI: 10.11646/zootaxa.4422.4.5 Reference page.
Benmesbah, M., Zacca, T., Casagrande, M.M., Mielke, O.H.H., Lamas, G. & Willmott, K.R. 2018. Taxonomic notes on Papilio ocypete Fabricius, 1776 and Papilio helle Cramer, 1779 with description of two new similar species from South America (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa 4425(1): 115–145. DOI: 10.11646/zootaxa.4425.1.7. Reference page.
Espeland, M., Breinholt, J.W., Willmott, K.R., Warren, A.D., Vila, R., Toussaint, E.F.A., Maunsell, S.C., Aduse-Poku, K., Talavera, G., Eastwood, R.G., Jarzyna, M.A., Guralnick, R., Lohman, D.J., Pierce, N.E. & Kawahara, A.Y. 2018. A Comprehensive and Dated Phylogenomic Analysis of Butterflies. Current Biology 28(5): 770–778. DOI: 10.1016/j.cub.2018.01.061 Reference page.
Nakahara, S., Zacca, T., Huertas, B., Neild, A.F.E., Hall, J.P.W., Lamas, G., Holian, L.A., Espeland, M. & Willmott, K.R. 2018: Remarkable sexual dimorphism, rarity and cryptic species: a revision of the ‘aegrota species group’ of the Neotropical butterfly genus Caeruleuptychia Forster, 1964 with the description of three new species (Lepidoptera, Nymphalidae, Satyrinae). Insect Systematics & Evolution 49(2): 130–182. DOI: 10.1163/1876312X-00002167 . Reference page.
Nakahara, S., Willmott, K.R., Mielke, O.H.H., Schwartz, J., Zacca, T., Espeland, M. & Lamas, G. 2018a: Seven new taxa from the butterfly subtribe Euptychiina (Lepidoptera: Nymphalidae: Satyrinae) with revisional notes on Harjesia Forster, 1964 and Pseudeuptychia Forster, 1964. Insecta Mundi 0639: 1–38. Reference page.
Willmott, K.R., Lamas, G., Radford, J.T., Marín, M.A., Nakahara, S., Espeland, M., Xiao, L. & Hall, J.P.W. 2018. A distinctive new species of cloud forest Euptychiina (Lepidoptera: Nymphalidae: Satyrinae) from Ecuador and Peru. Tropical Lepidoptera Research 28(1): 39-45. full article (PDF). Reference page.

2019

Freitas, Woodbury, M.S., Barbosa & Willmott 2019. A new subspecies of Adelpha messana (C. Felder & R. Felder) from the Brazilian Atlantic forest (Nymphalidae: Limenitidinae). Tropical Lepidoptera Research 29(2): 62-66. Online. Reference page. .
Nakahara, S., Lamas, G., Tyler, S., Marín, M.A., Huertas, B., Willmott, K.R., Mielke, O.H.H. & Espeland, M. 2019: A revision of the new genus Amiga Nakahara, Willmott & Espeland, gen. n., described for Papilio arnaca Fabricius, 1776 (Lepidoptera, Nymphalidae, Satyrinae). ZooKeys 821: 85–152. Reference page.
Willmott, K.R., Marín, M.A., Nakahara, S., Pomerantz, T., Lamas, G., Huertas, B., Espeland, M., Xiao, L., Hall, J.P.W., Willmott, J.I.R. & Freitas, A.V.L. 2019. A revision of the new Andean butterfly genus Optimandes Marín, Nakahara & Willmott, n. gen., with the description of a new species (Nymphalidae: Satyrinae: Euptychiina). Tropical Lepidoptera Research 29(1): 29-44. Online. Reference page. .

2020

Nakahara, S., Kleckner, K., Lamas, G., Huertas, B. & Willmott, K.R. 2020. A contribution towards the systematics of Magneuptychia Forster, 1964: Caeruleuptychia francisca (Butler, 1870), n. comb. (Lepidoptera: Nymphalidae: Satyrinae). Tijdschrift voor Entomologie 163(1): 51-61. Reference page.
Willmott, K.R., Lamas, G., Hall, J.P.W., Mota, L.L. & Kell, T. 2020. The common, the rare, and the lost: Descriptions of twelve new species and three new subspecies of equatorial Ithomiini (Lepidoptera, Nymphalidae, Danainae). Tropical Lepidoptera Research 30(1): 1-49. Reference page.
Zacca, T., Casagrande, M.M., Mielke, O.H.H., Huertas, B., Espeland, M., Freitas, A.V.L., Willmott, K.R., Nakahara, S. & Lamas, G. 2020. Revalidation of Vareuptychia Forster, 1964, description of Vanima gen. nov., and notes on Euptychia cleophes Godman & Salvin, 1889 (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa 4858(1): 1–34. DOI: 10.11646/zootaxa.4858.1.1. . Reference page.
Zacca, T., Casagrande, M.M., Mielke, O.H.H., Huertas, B., Barbosa, E.P., Freitas, A.V.L. & Willmott, K.R. 2020a. Description of Emeryus Zacca, Mielke & Casagrande gen. nov. (Lepidoptera: Nymphalidae) to accommodate three species formerly placed in Paryphthimoides Forster, 1964. Austral Entomology 59: 505–523. DOI: 10.1111/aen.12479. Reference page.

2021

Mahecha-Jiménez, O., Florczyk, K., Willmott, K.R., Cerdeña, J.A., Zubek, Boyer, P., Farfán, J., Lachowska-Cierlik, D. & Pyrcz, T.W. 2021. Solving the cryptic diversity of the genus Manerebia Staudinger in northern Peru: description of new species and considerations on the biogeographical role of the Huancabamba Deflection (Nymphalidae: Satyrinae: Pronophilina). Zootaxa 5072(3): 201–237. DOI: 10.11646/zootaxa.5072.3.1. . Reference page.
Nakahara, S., Matos-Maraví, P., Schwartz, J. & Willmott, K.R. 2021. Assessing a generic synapomorphy of Pseudodebis Forster, 1964 (Lepidoptera : Nymphalidae : Satyrinae) and a recent speciation with a shift in elevation between two new species in the western Andes. Invertebrate Systematics 35: 158–180. DOI: 10.1071/IS20024. . Reference page.
Nakahara, S., Matos-Maraví, P., Willmott, K.R., Nakamura, I. & MacDonald, J.R. 2021. Description of a new species of Pseudodebis Forster, 1964 from Central America (Lepidoptera: Nymphalidae: Satyrinae). Tropical Lepidoptera Research 31(1): 35-41. Reference page.
Nakahara, S., Willmott, K.R., MacDonald, J.R. & Thurman, A. 2021b. Description of an enigmatic new species of Chloreuptychia Forster, 1964 from Panama (Lepidoptera: Nymphalidae: Satyrinae). Tropical Lepidoptera Research 31(3): 145-150. Reference page.
Neild, A.F.E., Losada, M.E. & Willmott, K.R. 2021. A distinctive new subspecies of Moschoneura pinthous (Linnaeus, 1758) (Lepidoptera: Pieridae: Dismorphiinae) from the Orinoco Delta, Venezuela, with comments on the species-level taxonomy. Tropical Lepidoptera Research 31(1): 60-67. Reference page.
Padrón, P.S., Pyrcz, T.W. & Willmott, K.R. 2021. A Contribution Towards Resolving the Systematics of the High-Altitude Tropical Andean Satyrine Genus Altopedaliodes Forster, 1964 (Lepidoptera, Nymphalinae: Satyrinae). Neotropical Entomology 50: 767–803. DOI: 10.1007/s13744-021-00897-9. . Reference page.
Rosa, A.H.B., Huertas, B., Willmott, K.R., Barbosa, E.P., Machado, P.A., Mielke, O.H.H., Canaan, C.H.P. & Freitas, A.V.L. 2021. Fifty years without a name: a new species of Splendeuptychia Forster (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa 5061(1): 095–114. DOI: 10.11646/zootaxa.5061.1.4. . Reference page.
Willmott, K.R., Lamas, G., Hall, J.P.W., Vitale, F., Boyer, P., Petit, J.C., Radford, J.T. & Elias, M. 2021. A new species and thirty-eight new subspecies of equatorial Ithomiini (Lepidoptera, Nymphalidae, Danainae). Tropical Lepidoptera Research 31 (Supplement 3): 1-80. DOI: 10.5281/zenodo.5526837. Reference page.
Zacca, T., Casagrande, M.M., Mielke, O.H.H., Huertas, B., Barbosa, E.P., Freitas, A.V.L.. Lamas, G., Espeland, M., Brévignon, C., Nakahara, S., Checa, M.F. & Willmott, K.R. 2021. Systematics of the Neotropical butterfly genus Paryphthimoides Forster, 1964 (Lepidoptera: Nymphalidae: Satyrinae), with descriptions of seven new taxa. Insect Systematics & Evolution 52(1): 42–96. DOI: 10.1163/1876312X-00001027 . Reference page.
Zacca, T., Casagrande, M.M., Mielke, O.H.H., Huertas, B., Freitas, A.V.L.. Marín, M.A., Espeland, M. & Willmott, K.R. 2021a. A new euptychiine butterfly species from south Brazil and taxonomic rearrangements for Taydebis Freitas, 2013 and Hermeuptychia Forster, 1964 (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa 5023(4): 555–570. DOI: 10.11646/zootaxa.5023.4.5. . Reference page.

2022

Mota, L.L., Rosa, A.H.B., Vasconcellos, L.R., Willmott, K.R. & Freitas, A.V.L. 2022. A new subspecies of Mechanitis lysimnia from southern Amazonia (Nymphalidae: Danainae: Ithomiini). Tropical Lepidoptera Research 32(1): 47-51. Reference page.

2023

Hall, J.P.W. & Willmott, K.R. 2023. A review of the Argyrogrammana fauna of the Transandean region (Lepidoptera: Riodinidae: Symmachiini). Tropical Lepidoptera Research 33(supplement 1): 28-48. Reference page.
Hall, J.P.W., Willmott, K.R. & Ahrenholz, D.H. 2023. A review of the Argyrogrammana fauna of the eastern Andes (Lepidoptera: Riodinidae: Symmachiini) Tropical Lepidoptera Research 33(supplement 1): 49-60. Reference page.
Hall, J.P.W., Willmott, K.R. & Ahrenholz, D.H. 2023a. A review of the Argyrogrammana amalfreda group fauna of the western Amazon (Lepidoptera: Riodinidae: Symmachiini). Tropical Lepidoptera Research 33(supplement 1): 61-85. Reference page.
Nakahara, S., Kleckner, K., Barbosa, E.P., Lourenço, G.M., Casagrande, M.M., Willmott, K. & Freitas, A.V.L. 2023. Reassessment of the type locality of Euptychia stigmatica Godman, 1905, with the description of two new sibling species from Amazonia (Lepidoptera, Nymphalidae, Satyrinae, Satyrini). Zookeys 1167ː 57–88. DOI: 10.3897/zookeys.1167.102979 Reference page.

2024

Pyrcz, T.W., Boyer, P., Petit, J.C., Garlacz, R., Zajac-Garlacz, K.S., Espeland, M. & Willmott, K.R. 2024. Bedazzled: a new, striking species of Corades from the outskirts of Quito questions our knowledge of Andean cloud forest butterflies (Lepidoptera, Nymphalidae, Satyrinae). Zootaxa 5453(2): 255–262. DOI: 10.11646/zootaxa.5453.2.6. Reference page.
Willmott, K.R., Lamas, G., Hall, J.P.W., Boyer, P., Pyrcz, T.W. & Florczyk, K. 2024. A new species and eight new subspecies of high elevation Actinote from Colombia, Ecuador and Peru (Nymphalidae, Heliconiinae, Acraeini). Tropical Lepidoptera Research 34(1): 1-20. Reference page.

Links

List of publications. 2016. Systematics of Neotropical Butterflies. McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History.